1. Center of Ocean Mega-Science, Chinese Academy of Sciences, Qingdao, PR China
2. College of Earth Science, University of Chinese Academy of Sciences, Beijing 100049, PR China
3. Laboratory for Marine Biology and Biotechnology, Qingdao National Laboratory for Marine Science and Technology, Qingdao 266071, PR China
4. Key Laboratory of Experimental Marine Biology, Institute of Oceanology, Chinese Academy of Sciences, Qingdao 266071, PR China