1. School of Ecology, Sun Yat-Sen University, Guangzhou 510275, PR China
2. State Key Laboratory of Biocontrol, Guangdong Key Laboratory of Plant Resources and Southern Marine Science and Engineering Guangdong Laboratory (Zhuhai), School of Life Sciences, Sun Yat-Sen University, Guangzhou 510275, PR China
3. International Joint Research Center for Karstology, School of Earth Sciences, Yunnan University, Kunming 650091, PR China
4. State Key Laboratory of Desert and Oasis Ecology, Xinjiang Institute of Ecology and Geography, Chinese Academy of Sciences,, Urumqi 830011, PR China