1. Faculty of Science, University of the Ryukyus, 1 Senbaru, Nishihara, Okinawa 903-0213, Japan
2. Sakaerat Environmental Research Station, Thailand Institute of Scientific and Technological Research, 1 Moo 9 Udom, Wang Nam Khieo, Nakhon Ratchasima 30370, Thailand
3. Department of Marine Science, Kasetsart University, 50 Ngam Wong Wan Rd, Lat Yao Chatuchak, Bangkok 10900, Thailand