1. Australian School of Environmental Studies, Griffith University, Nathan, Qld 4111, Australia.
2. School of Environmental and Information Sciences, Charles Sturt University, PO Box 789, Albury, NSW 2640, Australia.
3. Royal Netherlands Institute for Sea Research (NIOZ), PO Box 59, 1790 AB Den Burg, Texel, The Netherlands.
4. Centre for Ecological and Evolutionary Studies, University of Groningen, PO Box 45, 9750 AA Haren, The Netherlands.