1. Research Center for Endangered Species, National Institute of Ecology, Yeongyang, Gyeongbuk Province 36531, Republic of Korea
2. Department of Biology, Kyung Hee University, Seoul 02447, Republic of Korea
3. College of Veterinary Medicine, Chungnam National University, Daejeon 34134, Republic of Korea
4. Department of Agriculture, Forestry, and Bioresources, Seoul National University, Seoul 08826, Republic of Korea