1. Institute of Wildlife Conservation, College of Veterinary Medicine, National Pingtung University of Science and Technology, Pingtung 912, Taiwan
2. Institute of Wildlife Conservation, College of Veterinary Medicine, National Pingtung University of Science and Technology, Pingtung 912, Taiwan and
3. Department of Biological Sciences, National Sun Yat-sen University, Gushan District, Kaohsiung City, 804, Taiwan