1. Department of Mathematical Sciences, Norwegian University of Science and Technology, N-7491 Trondheim, Norway;
2. Department of Mathematics and Statistics, The Arctic University of Norway, 9037 Tromsø, Norway
3. Centre for Research into Ecological and Environmental Modelling, School of Mathematics and Statistics, University of St. Andrews, KY16 9LZ Fife, United Kingdom
4. Department of Mathematical Sciences, University of Bath, BA2 7AY Bath, United Kingdom
5. School of Mathematics, The University of Edinburgh, EH9 3FD Edinburgh, United Kingdom