1. VIB Center for the Biology of Disease, Vlaams Instituut voor Biotechnologie, Leuven 3000, Belgium;,
2. Center for Human Genetics, KU Leuven, Leuven 3000, Belgium
3. Leuven Research Institute for Neuroscience & Disease (LIND), KU Leuven, Leuven 3000, Belgium
4. Institute of Neurology, University College London, WC1N 3BG London, United Kingdom