1. Division of Mucosal Immunology, Department of Microbiology and Immunology, The Institute of Medical Science, The University of Tokyo, Tokyo 108-8639, Japan;
2. International Research and Development Center for Mucosal Vaccines, The Institute of Medical Science, The University of Tokyo, Tokyo 108-8639, Japan
3. Institute for Global Prominent Research, Chiba University, Chiba 260-8670, Japan
4. Department of Mucosal Immunology, Graduate School of Medicine, Chiba University, Chiba 260-8670, Japan
5. Department of Innovative Medicine, Graduate School of Medicine, Chiba University, Chiba 260-8670, Japan
6. Chiba University–UC San Diego Center for Mucosal Immunology, Allergy, and Vaccine, La Jolla, CA 92093
7. Department of Immunology, Graduate School of Medicine, Chiba University, Chiba 260-8670, Japan