1. CIRAD, UPR Bioagresseurs, 30501 Turrialba, Costa Rica; Bioagresseurs, Université de Montpellier, CIRAD, 34090 Montpellier, France;
2. Program of Sustainable Agriculture and Agroforestry, Tropical Agricultural Research and Higher Education Center (CATIE), 30501 Turrialba, Costa Rica;, ,
3. Inter-American Institute for Cooperation on Agriculture (IICA), 11101 Coronado, San José, Costa Rica
4. CIRAD, UMR SYSTEM, 30501 Turrialba, Costa Rica; SYSTEM, Université de Montpellier, CIHEAM-IAMM, CIRAD, INRA, 34090 Montpellier SupAgro, Montpellier, France;
5. UMR AGIR, Institut National de la Recherche Agronomique (INRA), Université de Toulouse, INPT, INP-EI Purpan, Castanet-Tolosan, France;,