1. School of GeoSciences, University of Edinburgh, Edinburgh EH8 9XP, United Kingdom;
2. Institute of Biological and Environmental Sciences, School of Biological Sciences, University of Aberdeen, Aberdeen AB24 3UU, United Kingdom;
3. Department of Bioscience, Aarhus University, 4000 Roskilde, Denmark
4. Department of Physical Geography and Ecosystem Science, Lund University, 221 00 Lund, Sweden;
5. Ocean and Earth Science, National Oceanography Centre Southampton, University of Southampton, Southampton SO17 1BJ, United Kingdom;
6. Grasslands Research Centre, New Zealand Agricultural Greenhouse Gas Research Centre, Palmerston North 4442, New Zealand;