1. College of Science, Health, Engineering and Education, Murdoch University, Australia; e-mail: jessica.rendle@twycrosszoo.org, l.yeap@murdoch.edu.au, b.jackson@murdoch.edu.au, k.warren@murdoch.edu.au, r.donaldson@murdoch.edu.au, j.richardson@murdoch.e
2. School of Animal, Rural and Environmental Sciences, Nottingham Trent University, UK; e-mail: samantha.ward@ntu.ac.uk
3. School of Animal Biology, University of Western Australia, Australia; e-mail: c.mayberry@ymail.com