1. Department of Fish and Wildlife Conservation, Virginia Tech, Cheatham Hall, 310 W. Campus Drive, Blacksburg, Virginia 24061, U.S.A.
2. The Nature Conservancy in Virginia, Volgenau Virginia Coast Reserve, 11332 Brownsville Road – PO Box 158, Nassawadox, Virginia 23413, U.S.A.
3. Virginia Department of Wildlife Resources, 12610 Jacobus Creek Road, Machipongo, Virginia 23405, U.S.A.
4. Department of Behavioural Ecology & Evolutionary Genetics, Max Planck Institute for Ornithology, Eberhard Gwinner Str, Seewiesen 82319, Germany