1. Department of Genetics, Washington University School of Medicine, St. Louis, MO 63108, USA.
2. Edison Family Center for Genome Sciences and Systems Biology, Washington University School of Medicine, St. Louis, MO 63108, USA.
3. Maryville University of St Louis, St. Louis, MO 63141, USA.
4. Department of Developmental Biology, Washington University School of Medicine, St. Louis, MO 63108, USA.
5. Department of Neuroscience, Washington University School of Medicine, St. Louis, MO 63108, USA.