1. Institute of Zoology, Zoological Society of London, Regent’s Park, London NW1 4RY, UK.
2. Ocean and Earth Science, National Oceanography Centre Southampton, University of Southampton, Waterfront Campus, European Way, Southampton SO14 3ZH, UK.
3. Department of Anthropology, Stony Brook University, Stony Brook, NY 11794, USA.
4. Centre ValBio, Ranomafana, Ifanadiana 312, Madagascar.
5. Mention Bassins Sédimentaires Evolution Conservation, University of Antananarivo, Antananarivo, Madagascar.
6. Department of Anthropology, University of Massachusetts, 240 Hicks Way, Amherst, MA 01003, USA.
7. School of Science, Engineering and Design, Teesside University, Borough Road, Middlesbrough TS1 3BA, UK.