1. Centro de Biología Molecular Severo Ochoa, Consejo Superior de Investigaciones Científicas–Universidad Autónoma de Madrid, Madrid 28049, Spain.
2. British Antarctic Survey, Natural Environment Research Council, Cambridge CB3 0ET, UK.
3. Faculty of Health and Life Sciences, University of Northumbria, Newcastle upon Tyne NE1 8ST, UK.
4. University Centre in Svalbard, Longyearbyen N-9171, Norway.