1. Fenner School of Environment and Society, Australian National University, Canberra, Australian Capital Territory 0200, Australia.
2. School of Biological Sciences, University of Queensland, St. Lucia, Queensland 4072, Australia.
3. Centre for Integrative Ecology, School of Life and Environmental Sciences, Deakin University, Geelong, Victoria 3216, Australia.
4. Research School of Biology, Australian National University, Canberra, Australian Capital Territory 0200, Australia.