1. Kobe University Research Center for Inland Seas, Awaji, Hyogo 656-2401, Japan.
2. Kobe University Graduate School of Science and Technology, Kobe, Hyogo 657-8501, Japan.
3. Department of Technology and Ecology, Hall of Global Environmental Research, Kyoto University, Kyoto 606-8501, Japan.
4. Graduate School of Human and Environmental Studies, Kyoto University, Kyoto 606-8501, Japan.
5. PRESTO, Japan Science and Technology Agency, Kawaguchi, Saitama, 332-0012, Japan.