1. Department of Biology, McGill University, Montréal, Québec H3A 1B1, Canada.
2. Department of Integrative Biology, University of Guelph, Guelph, Ontario N1G 2W1, Canada.
3. Perimeter Institute for Theoretical Physics, Waterloo, Ontario N2L 2Y5, Canada.
4. Biodiversity Research Centre, University of British Columbia, Vancouver, British Columbia V6T 1Z4, Canada.
5. Tanzania Wildlife Research Institute, P.O. Box 661, Arusha, United Republic of Tanzania.
6. Centre for Biodiversity Theory and Modeling, Experimental Ecology Station, CNRS, 09200 Moulis, France.