1. Institute for Regenerative Medicine, Perelman School of Medicine, University of Pennsylvania, Philadelphia, PA 19104-5157, USA.
2. Department of Cell and Developmental Biology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, PA 19104-5157, USA.
3. Department of Biochemistry and Molecular Biology and Tulane Center for Aging, Tulane University Health Sciences Center, New Orleans, LA 70112, USA.
4. The Institute for Translational Medicine and Therapeutics, Department of Genetics, Perelman School of Medicine, University of Pennsylvania, Philadelphia, PA 19104, USA.
5. Department of Bioengineering, University of Pennsylvania, Philadelphia, PA 19104, USA.
6. Howard Hughes Medical Institute, Department of Pharmacology, University of Texas Southwestern Medical Center, Dallas, TX 75390, USA.