1. Whitehead Institute for Biomedical Research and Department of Biology, Massachusetts Institute of Technology, Cambridge, MA 02142, USA.
2. Howard Hughes Medical Institute, Department of Biology, Massachusetts Institute of Technology, Cambridge, MA 02139, USA.
3. Koch Institute for Integrative Cancer Research, Massachusetts Institute of Technology, Cambridge, MA 02142, USA.
4. Broad Institute of Harvard and MIT, Cambridge, MA 02142, USA.
5. Leibniz Institute on Aging–Fritz Lipmann Institute, 07745 Jena, Germany.