1. Broad Institute of MIT and Harvard, Cambridge, MA 02142, USA.
2. Big Data Institute, Li Ka Shing Centre for Health Information and Discovery, University of Oxford, Oxford OX3 7LF, UK.
3. Department of Human Evolutionary Biology, Harvard University, Cambridge, MA 02138, USA.
4. Department of Genetics, Harvard Medical School, Boston, MA 02115, USA.
5. Howard Hughes Medical Institute, Harvard Medical School, Boston, MA 02115, USA.
6. Department of Evolutionary Anthropology, University of Vienna, 1090 Vienna, Austria.