1. Department of Molecular Biology, Princeton University, Washington Road, Princeton, NJ 08544, USA.
2. Program in Developmental & Stem Cell Biology, The Hospital for Sick Children, 686 Bay Street, Toronto, Ontario M5G 0A4, Canada.
3. Department of Molecular Genetics, The University of Toronto, Toronto, Ontario M5S 1A8, Canada.
4. Mouse Imaging Centre, The Hospital for Sick Children, 25 Orde Street, Toronto, Ontario M5T 3H7, Canada.
5. Department of Medical Biophysics, The University of Toronto, Toronto, Ontario M5G 2M9, Canada.