1. U.S. Geological Survey Biological Resources Division, Western Ecological Research Center, Sequoia–Kings Canyon Field Station, 47050 Generals Highway, Three Rivers, CA 93271–9651, USA.
2. Center for Environmental Analysis–Centers for Research Excellence in Science and Technology, Department of Biology and Microbiology, California State University, Los Angeles, CA 90032, USA.
3. U.S. National Park Service, Santa Monica Mountains National Recreation Area, Thousand Oaks, CA 91360, USA.