1. Department of Biology, Dalhousie University, Halifax, Nova Scotia, Canada B3H 4J1; .
2. Department of Ecology and Evolutionary Biology, Princeton University, Princeton, NJ 08544, USA; .
3. Department of Biology, University of California-San Diego, La Jolla, CA 92093, USA; .
4. Department of Biological Science, Florida State University, Tallahassee, FL 32306, USA; .
5. Department of Ecology, Evolution, and Marine Biology, University of California, Santa Barbara, CA 93106, USA; .
6. Department of Biology, University of Washington, Seattle, WA 98195, USA;