1. Department of Biology, University of Utah, 257 S. 1400 East, Salt Lake City, UT 84112, USA.
2. Environmental and Conservation Programs and Department of Botany, Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, USA.
3. Department of Forestry, Universidad Nacional de la Amazonía Peruana, Plaza Serafín Filomeno 246, Iquitos, Peru.