1. Key Laboratory of Ecology of Rare and Endangered Species and Environmental Protection, Guangxi Normal University, Ministry of Education, Guilin, Guangxi, China.
2. Guangxi Key Laboratory of Rare and Endangered Animal Ecology, College of Life Sciences, Guangxi Normal University, Guilin, Guangxi, China.
3. The Chongzuo White-Headed Langur Field Observation and Research Station of Guangxi, Chongzuo, Guangxi, China.
4. School of Life Sciences, University of KwaZulu-Natal, Scottsville, South Africa.
5. Institute of Biodiversity and Environmental Conservation, Universiti Malaysia Sarawak, Kota Samarahan, Sarawak, Malaysia.