1. Department of Ecology and Evolutionary Biology, Princeton University, Princeton, NJ 08544, USA.
2. Department of Ecology, Evolution, and Natural Resources and the Institute of Marine and Coastal Sciences, Rutgers University, New Brunswick, NJ 08901, USA.
3. Department of Biology, Dalhousie University, Halifax, NS B3H 4R2, Canada.
4. Northeast Fisheries Science Center, Woods Hole, MA 02453, USA.
5. Atmospheric and Oceanic Sciences Program, Princeton University, Princeton, NJ 08544, USA.