1. Department of Ecology and Evolution, University of Chicago, 1101 East 57th Street, Chicago, IL 60637, USA.
2. Laboratoire Génétique et Evolution des Populations Végétales, FRE CNRS 3268, Université des Sciences et Technologies de Lille 1, Villeneuve d'Ascq, France.
3. Department of Plant Pathology, Kansas State University, Manhattan, KS 66502, USA.