1. Molecular Biology Program, Memorial Sloan-Kettering Cancer Center, New York, NY 10065, USA.
2. Developmental Biology Program, Memorial Sloan-Kettering Cancer Center, New York, NY 10065, USA.
3. Department of Public Health and Cell Biology, Section of Anatomy, University of Rome Tor Vergata, 00133 Rome, Italy.
4. Howard Hughes Medical Institute, Memorial Sloan-Kettering Cancer Center, New York, NY 10065, USA.