1. Moss Landing Marine Laboratories, Moss Landing, CA 95039, USA
2. Department of Biology, San Diego State University, San Diego, CA 92182, USA
3. Marine Science Institute, University of California, Santa Barbara, CA 93106-6051, USA
4. Marine Ecology Research Group, School of Biological Sciences, University of Canterbury, Christchurch, New Zealand
5. Department of Ocean, Earth and Atmospheric Sciences, Old Dominion University, Norfolk, VA 23529, USA