1. Department of Chemical and Biological Engineering, University of Wisconsin–Madison, 1415 Engineering Drive, Madison, WI 53706–1607, USA.
2. Department of Biomolecular Chemistry, University of Wisconsin–Madison, 1300 University Avenue, Madison, WI 53706–1510, USA.
3. Department of Ophthalmology and Vision Science, School of Medicine, University of California, Davis, CA 95616, USA.
4. Department of Veterinary Surgical and Radiological Sciences, School of Veterinary Medicine, University of California, Davis, CA 95616, USA.