1. Department of Fisheries and Wildlife, Oregon State University, Corvallis, OR 97331–3803, USA.
2. Department of Vertebrate Zoology, National Museum of Natural History, Smithsonian Institution, Washington, DC 20013–7012, USA.
3. Center for the Study of Biological Diversity, University of Guyana, Turkeyen Campus, East Coast Demerara, Guyana.
4. Department of Ecology and Evolutionary Biology and Department of Biological Sciences, University of Toronto at Scarborough, Toronto, ON, M1C 1A4, Canada.