1. Department of Organismic and Evolutionary Biology, Harvard University, 16 Divinity Avenue, Cambridge, MA 02138, USA.
2. Environment Department, University of York, York, YO10 5DD, UK.
3. Southeast Fisheries Science Center, National Oceanic and Atmospheric Administration Fisheries, 75 Virginia Beach Drive, Miami, FL 33149, USA.
4. Department of Marine Resource and Environmental Management, Faculty of Natural Science, University of the West Indies, Cave Hill, Barbados.