1. Whitehead Institute for Biomedical Research and Department of Biology, Massachusetts Institute of Technology, Cambridge, MA 02142, USA.
2. Department of Pathology, Massachusetts General Hospital, Boston, MA 02114, USA.
3. Department of Pathology, Tufts University Schools of Medicine and Veterinary Medicine, Boston, MA 0211 1, USA.
4. Division of Pulmonary Biology, Children's Hospital, Cincinnati, OH 45229, USA.