1. Department of Ecology and Evolutionary Biology, University of Colorado, Boulder, CO 80309, USA.
2. Cooperative Institute for Research in Environmental Sciences, University of Colorado, Boulder, CO 80309, USA.
3. The Gladstone Institutes, University of California, San Francisco, CA 94158, USA.
4. Department of Genetics and Genomic Sciences and Department of Medicine, Mount Sinai School of Medicine, New York, NY 10029, USA.
5. Institute of Genomic and Systems Biology, Argonne National Laboratory, Argonne, IL 60439, USA.
6. Computation Institute, University of Chicago, Chicago, IL 60637, USA.
7. Institute for Human Genetics and Division of Biostatistics, University of California, San Francisco, CA 94143, USA.
8. Department of Chemistry and Biochemistry, University of Colorado, Boulder, CO 80309, USA.
9. Howard Hughes Medical Institute, Boulder, CO 80309, USA.
10. Department of Ecology and Evolution, University of Chicago, Chicago, IL 60637, USA.
11. Department of Plant and Soil Sciences, University of Kentucky, Lexington, KY 40546, USA.