1. Department of Ecology & Evolutionary Biology, The University of Connecticut, 1 University Place, Stamford, CT 06901-2315, USA
2. Department of Ecology and Evolutionary Biology, The University of Connecticut, 75 N Eagleville Road, Storrs, CT 06269-3043, USA
3. Department of Marine Sciences, The University of Connecticut, 1080 Shennecossett Road, Groton, CT 06340, USA
4. Department of Biological Sciences, The University of New Hampshire, 46 College Road, Durham, NH 03824, USA