1. Institute of Marine Biology/Center of Excellence for Marine Bioenvironment and Biotechnology, National Taiwan Ocean University, Keelung 20224, Taiwan, Republic of China
2. National Institute for Water and Atmospheric Research (NIWA), Private Bag 14-901, Wellington 6241, New Zealand
3. Department of Biology, University of North Carolina at Chapel Hill, Chapel Hill, North Carolina 27599-3280, USA