1. Guangzhou Key Laboratory of Subtropical Biodiversity and Biomonitoring, Guangdong Provincial Key Laboratory of Healthy and Safe Aquaculture, College of Life Science, South China Normal University, Guangzhou 510631, P. R. China
2. The Eighth Geological Brigade, Hebei Geological Prospecting Bureau, Qinhuangdao 066001, P. R. China
3. Natural History Museum of Denmark, University of Copenhagen, Sølvgade 83S, Copenhagen, Denmark