1. Seaweed and Seagrass Research Unit, Excellence Center for Biodiversity of Peninsula Thailand, Department of Biology, Faculty of Science, Prince of Songkla University, Hat Yai, Songkhla 90112, Thailand
2. School of Biological Sciences, Victoria University of Wellington, P.O. Box 600, Wellington, 6140, New Zealand
3. Institute of Ocean and Earth Sciences, University of Malaya, Kuala Lumpur, 50603 Malaysia
4. Research Center for Inland Seas, Kobe University, Rokkodai, Kobe 657-8501, Japan