1. Kobe University Research Center for Inland Seas, Kobe 657-8501, Japan
2. Institute for International Collaboration, Hokkaido University, Sapporo 060-0808 Japan
3. Senckenberg am Meer, German Centre for Marine Biodiversity Research (DZMB), Südstrand 44, D-26382 Wilhelmshaven, Germany
4. Faculty of Science, Hokkaido University, Sapporo 060-0810 Japan