1. Faculty of Sciences of Tunis, Biology Depart- ment, Research Unit of Physiology and Aquatic Environment, University of Tunis El Manar, 2092 Tunis, Tunisia
2. Aquatic Environment Exploitation Resources Unit, Higher institute fishing and fish farming of Bizerte, Tunisia.
3. Faculty of Sciences of Tunis, Biology Depart- ment, Research Unit of Physiology and Aquatic Environment, University of Tunis El Manar, 2092 Tunis, Tunisia.