1. Shanghai Key Laboratory of Regulatory Biology, Joint Research Center for Translational Medicine, ECNU-Fengxian Hospital, Institute of Biomedical Sciences and School of Life Sciences, East China Normal University, Shanghai 200241, China
2. Fengxian Hospital affiliated to Southern Medical University, Shanghai, 201499, China
3. Life Sciences Institute, Zhejiang University, Hangzhou 310058, China
4. Hubrecht Institute, Royal Netherlands Academy of Arts and Sciences (KNAW), Uppsalalaan 8, 3584 CT, Utrecht the Netherlands