1. Warwick Medical School, University of Warwick 1 Division of Biomedical Sciences , , Coventry, CV2 2DX , UK
2. Tommy's National Centre for Miscarriage Research, University Hospitals Coventry & Warwickshire NHS Trust 2 , Warwick Medical School, University of Warwick, Coventry, CV2 2DX , UK
3. University at Buffalo 3 Department of Biological Sciences , , Buffalo, NY 14260-4610 , USA
4. Johns Hopkins University 4 Department of Biology , , Baltimore, MD 21218 , USA