1. Centre d'Etudes Biologiques de Chizé, CNRS, La Rochelle Université, UMR 7372 1 , Villiers-en-Bois , France
2. Lund University 2 Department of Biology , , Ecology Building, 223 62 Lund , Sweden
3. University of Southern Denmark 3 Department of Biology , , 5230 Odense , Denmark