1. Department of General Systems Studies, Graduate School of Arts and Sciences, The University of Tokyo, Komaba, Tokyo 3-8-1, Japan
2. Laboratory of Ecological Genetics, Graduate School of Environmental Science, Hokkaido University, Sapporo 060-0810, Japan
3. Department of Bioresource Science, Tamagawa University, Machida, Tokyo 194-8610, Japan
4. Department of Hygiene and Public Health, Tokyo Women's Medical University, Shinjuku-ku, Tokyo 162-8666, Japan
5. Department of Subtropical Agro-Environmental Sciences, Faculty of Agriculture, University of the Ryukyus, Nishihara, Okinawa 903-0213, Japan