1. University of Louisville 1 Department of Biology , , Louisville, KY 40208 , USA
2. University of Louisville 2 Department of Anatomical Sciences & Neurobiology , , Louisville, KY 40202 , USA
3. Saint Louis University 3 Department of Biology , , St Louis, MO 63103 , USA
4. East Carolina University 4 Department of Biology , , Greenville, NC 27858
5. Hokkaido University 5 Faculty of Science , , Sapporo 060-0810 , Japan