1. Department of Biological Sciences, University of Essex, Colchester, Essex CO4 5AP, UK
2. School of Agricultural Science, University of Tasmania, Private Bag 54, Hobart, Tasmania 7001, Australia
3. Crop Performance and Improvement Division, Rothamsted Research, Harpenden, Hertfordshire AL5 2JQ, UK
4. Department of Physiology and Biochemistry of Plants, Biological Faculty, Belarusian State University, Minsk 220030, Belarus