1. Evolution & Ecology Research Centre, School of Biological, Earth and Environmental Sciences, UNSW Sydney, Sydney, NSW 2052, Australia
2. Division of Ecology and Evolution, Research School of Biology, The Australian National University, Canberra, ACT 2600, Australia
3. Department of Biology, Mount Allison University, Sackville, NB, Canada, E4L 1E2
4. Charles Perkins Centre, Faculty of Science, School of Life and Environmental Sciences and School of Mathematics and Statistics, The University of Sydney, Sydney, NSW 2006, Australia
5. Department of Biological Sciences, College of Sciences and Mathematics, Auburn University, Auburn, AL 36849, USA
6. School of Natural Sciences, University of Tasmania, Sandy Bay, Hobart, TAS 7001, Australia