1. Max Planck Research Group ‘Chromatin and Ageing’, Max Planck Institute for Biology of Ageing, Joseph-Stelzmann-Str. 9b, 50931 Cologne, Germany
2. Biozentrum, University of Basel, Klingelbergstrasse 50/70, 4056 Basel, Switzerland
3. Max Planck Research Group ‘Autophagy and Cellular Ageing‘, Max Planck Institute for Biology of Ageing, Joseph-Stelzmann-Str. 9b, 50931 Cologne, Germany
4. Cologne Excellence Cluster on Stress Responses in ageing-associated Diseases (CECAD), University of Cologne, Joseph-Stelzmann-Str. 26, 50931 Cologne, Germany